1980, PhD in Medical and Pphysiological Chemistry, University of Uppsala. Since 1986, Director, Ludwig Institute for Cancer Research, Uppsala. Professor of Molecular Cell Biology, Uppsala University. Research focuses on mechanisms whereby growth factors activate their receptors and control cell growth, migration, apoptosis and differentiation, and how perturbations of signalling pathways promote tumour progression. Former: Member, Board, Swedish Research Council; Vice-President, European Research Council. Chair of the Board: Nobel Foundation; Science for Life Laboratory; European Molecular Biology Organization. Associate Editor, Molecular Biology of the Cell, Genes to Cells and Growth Factors. Member, past or present, scientific advisory boards, including: European Molecular Biology Organization, Heidelberg; German Cancer Center, Heidelberg; Max Planck Institute for Biochemistry, Martinsried; Center for Molecular Medicine, Vienna; Finnish Institute for Molecular Medicine, Helsinki; Biocenter Finland; Institute for Cancer Research, Oslo; Center for Cancer Biomarkers, Bergen; K.G. Jebsen Foundation, Bergen. Elected Member: European Molecular Biology Organization; Royal Swedish Academy of Sciences; Royal Scientific Society; Academia Europea; Japanese Association for Cancer Research; Finnish Scientific Society; ScanBalt Academy; Hellenic Biochemical Society; Norwegian Academy of Sciences and Letters; European Academy of Cancer Sciences. Author of 418 research articles and 205 review articles; holder of 42 approved patents. Recipient of numerous honours and awards.