1960, BA (Hons) in Chemistry, Swarthmore College; 1964, PhD in Biology, Rockefeller Univ. Post-doctoral fellow, MIT and Albert Einstein College of Medicine; 1965-68, Research Assoc., Salk Institute for Biological Studies; 1968-72, Assoc. Professor of Microbiology, MIT; 1972-90, Professor of Biology; 1973-83, American Cancer Society Professor of Microbiology; 1982-90, Director, Whitehead Institute for Biomedical Research; 1990-91, President, Rockefeller Univ.; 1990-94, Professor, Rockefeller Univ.; 1994-97, Ivan R. Cottrel Professor of Molecular Biology and Immunology, MIT; 1994-97, American Cancer Society Research Professor; 1995-97, Institute Professor, MIT; 1996-2002, Chairman, NIH AIDS Vaccine Research Cttee; since 1997, President, California Institute of Technology. 1970, discovered reverse transcriptase with Dr Temin. 1974, National Academy of Arts and Sciences. 1978, Member, Pontifical Academy of Sciences. Foreign Member, Royal Society (England); 1988, Institute of Medicine. 1975, Nobel Prize in Physiology or Medicine. Recipient of the 1999 National Medal of Science and numerous other awards and honours.