1970, BA, Yale; 1976, PhD, Harvard. Former Professor of Neurobiology, Washington University, St Louis. Professor of Molecular and Cellular Biology and Director, Center for Brain Science, Harvard University. Member, US National Academy of Sciences and American Academy of Arts and Sciences. Basic research on the development of the nervous system, especially formation of specific patterns of connections among nerve cells in the visual system that underlie visual function. Also develops novel methods (e.g., Brainbow) to visualize neuronal circuits.